Allentoft, M. E. et al. Population genomics of Bronze Age Eurasia. Nature 522, 167–172 (2015).
Google Scholar
Haak, W. et al. Massive migration from the steppe was a source for Indo-European languages in Europe. Nature 522, 207–211 (2015).
Google Scholar
Allentoft, M. E. et al. Population genomics of post-glacial western Eurasia. Nature https://doi.org/10.1038/s41586-023-06865-0 (2024).
Posth, C. et al. Palaeogenomics of Upper Palaeolithic to Neolithic European hunter-gatherers. Nature 615, 117–126 (2023).
Google Scholar
Johannsen, N. N., Larson, G., Meltzer, D. J. & Vander Linden, M. A composite window into human history. Science 356, 1118–1120 (2017).
Google Scholar
Furholt, M. Mobility and social change: understanding the European Neolithic period after the archaeogenetic revolution. J. Archaeol. Res. 29, 481–535 (2021).
Google Scholar
Kristiansen, K. Archaeology and the Genetic Revolution in European Prehistory (Elements in the Archaeology of Europe) (Cambridge Univ. Press, 2022).
Fischer, A. & Kristiansen, K. The Neolithisation of Denmark. 150 Years of Debate (J. R. Collis, 2002).
Günther, T. et al. Population genomics of Mesolithic Scandinavia: investigating early postglacial migration routes and high-latitude adaptation. PLoS Biol. 16, e2003703 (2018).
Google Scholar
Kashuba, N. et al. Ancient DNA from mastics solidifies connection between material culture and genetics of mesolithic hunter–gatherers in Scandinavia. Commun. Biol. 2, 185 (2019).
Google Scholar
Fischer, A. in The Neolithisation of Denmark—150 years of debate (eds Fischer, A. & Kristiansen, K.) 343–393 (J. R. Collis, 2002).
Price, T. D. Europe’s First Farmers (Cambridge Univ. Press, 2000).
Lipson, M. et al. Parallel palaeogenomic transects reveal complex genetic history of early European farmers. Nature 551, 368–372 (2017).
Google Scholar
Mathieson, I. et al. The genomic history of southeastern Europe. Nature 555, 197–203 (2018).
Google Scholar
Brace, S. et al. Ancient genomes indicate population replacement in Early Neolithic Britain. Nat. Ecol. Evol. 3, 765–771 (2019).
Google Scholar
Midgley, M. TRB Culture: The First Farmers of the North European Plain (Edinburgh Univ. Press, 1992).
Iversen, R. in Tracing the Indo-Europeans: New evidence from Archaeology and Historical Linguistics (eds Olsen, B. A., Olander, T. & Kristiansen, K.) 73–95 (Oxbow, 2019).
Nielsen, S. K. & Johannsen, N. N. Mortuary palisades, single graves, and cultural admixture: the establishment of Corded Ware culture on the Jutland Peninsula. Praehistorische Zeitschrift https://doi.org/10.1515/pz-2023-2022 (2023).
Google Scholar
Kristiansen, K. Prehistoric migrations—the case of the Single Grave and Corded Ware Cultures. J. Dan. Archaeol. 8, 211–225 (1991).
Google Scholar
Lewis, J. P. et al. Marine resource abundance drove pre-agricultural population increase in Stone Age Scandinavia. Nat. Commun. 11, 2006 (2020).
Google Scholar
Sousa da Mota, B. et al. Imputation of ancient human genomes. Nat. Commun. 14, 3660 (2023).
Google Scholar
Schilling, H. in Mesolithic on the Move (eds Larsson, L. et al.) 351–358 (2003).
Petersen, P. V. Chronological and regional variation in the Late Mesolithic of Eastern Denmark. J. Dan. Archaeol. 3, 7–18 (1984).
Google Scholar
Fischer, A. in The Danish Storebælt Since the Ice Age (eds Pedersen, L., Fischer, A. & Aaby, B.) 63–77 (Sea and Forest, 1997).
Sørensen, S. A. The Kongemose Culture (Univ. Press of Southern Denmark, 2017).
Dolbunova, E. et al. The transmission of pottery technology among prehistoric European hunter-gatherers. Nat. Hum. Behav. 7, 171–183 (2023).
Google Scholar
Klassen, L. Jade und Kupfer: Untersuchungen zum Neolithisierungsprozess im westlichen Ostseeraum unter besonderer Berücksichtigung der Kulturentwicklung Europas 5500–3500 bc vol. 47 (Aarhus Universitetsforlag, 2004).
Price, T. D. Seeking the First Farmers in Western Sjælland, Denmark: The Archaeology of the Transition to Agriculture in Northern Europe (Oxbow Books, 2022).
Hansen, J. et al. The Maglemosian skeleton from Koelbjerg revisited: Identifying sex and provenance. Dan. J. Archaeol. 6, 55–66 (2017).
Sørensen, M. in Ecology of Early Settlement in Northern Europe: Conditions for Subsistence and Survival. The Early Settlement of Northern Europe Vol. 1 (eds. Persson, P., Riede, F. & Skar, B.) 277–301 (Equinox, 2018).
Piezonka, H. et al. Stone Age pottery chronology in the Northeast European Forest Zone: new AMS and EA-IRMS results on foodcrusts. Radiocarbon 58, 267–289 (2016).
Google Scholar
Mathieson, I. et al. Genome-wide patterns of selection in 230 ancient Eurasians. Nature 528, 499–503 (2015).
Google Scholar
Irving-Pease, E. K. et al. The selection landscape and genetic legacy of ancient Eurasians. Nature https://doi.org/10.1038/s41586-023-06705-1 (2024).
Fischer, A. et al. Coast-inland mobility and diet in the Danish Mesolithic and Neolithic: evidence from stable isotope values of humans and dogs. J. Archaeol. Sci. 34, 2125–2150 (2007).
Google Scholar
Fischer, A. et al. The composition of Mesolithic food—evidence from a submerged settlement on the Argus Bank, Denmark. Acta Archaeol. 78, 163–178 (2007).
Google Scholar
Brinch Petersen, E. Gravene ved Dragsholm. Fra jægere til bønder for 6000 år siden. Nationalmuseets Arbejdsmark 1974, 112–120 (1974).
Price, T. D. et al. New information on the Stone Age graves at Dragsholm, Denmark. Acta Archaeol. 78, 193–219 (2007).
Google Scholar
Sørensen, L. From Hunter to Farmer in Northern Europe. Migration and adaptation during the Neolithic and Bronze Age. Acta Archaeologica Vol. 85 (Wiley–Blackwell, 2014).
Nielsen, P. O. & Nielsen, F. O. S. First Farmers on the Island of Bornholm (The Royal Society of Northern Antiquaries and Univ. Press of Southern Denmark, 2020).
Sjögren, K.-G. & Fischer, A. The chronology of Danish dolmens. Results from 14C dates on human bones. J. Neolit. Archaeol. 25, https://doi.org/10.12766/jna.2023.1 (2023).
Dehn, T. & Hansen, S. I. Birch bark in Danish passage graves. J. Dan. Archaeol. 14, 23–44 (2006).
Google Scholar
Ebbesen, K. Simple, tidligneolitiske grave. Aarbøger for nordisk Oldkyndighed og Historie 1992, 47–102 (1994).
Gron, K. J. & Sørensen, L. Cultural and economic negotiation: a new perspective on the Neolithic Transition of Southern Scandinavia. Antiquity 92, 958–974 (2018).
Google Scholar
Chintalapati, M., Patterson, N. & Moorjani, P. The spatiotemporal patterns of major human admixture events during the European Holocene. eLife 11, e77625 (2022).
Google Scholar
González-Fortes, G. et al. Paleogenomic evidence for multi-generational mixing between Neolithic farmers and Mesolithic hunter-gatherers in the Lower Danube Basin. Curr. Biol. 27, 1801–1810.e10 (2017).
Google Scholar
Villalba-Mouco, V. et al. Survival of Late Pleistocene hunter-gatherer ancestry in the Iberian Peninsula. Curr. Biol. 29, 1169–1177.e7 (2019).
Google Scholar
Jensen, T. Z. T. et al. A 5700 year-old human genome and oral microbiome from chewed birch pitch. Nat. Commun. 10, 5520 (2019).
Google Scholar
Olalde, I. et al. Derived immune and ancestral pigmentation alleles in a 7,000-year-old Mesolithic European. Nature 507, 225–228 (2014).
Google Scholar
Cox, S. L., Ruff, C. B., Maier, R. M. & Mathieson, I. Genetic contributions to variation in human stature in prehistoric Europe. Proc. Natl Acad. Sci. USA 116, 21484–21492 (2019).
Google Scholar
Klassen, L. (ed.) The Pitted Ware Culture on Djursland: Supra-regional Significance and Contacts in the Middle Neolithic of Southern Scandinavia (Aarhus Univ. Press, 2020).
Iversen, R., Philippsen, B. & Persson, P. Reconsidering the Pitted Ware chronology. Praehistorische Zeitschrift 96, 44–88 (2021).
Google Scholar
Coutinho, A. et al. The Neolithic Pitted Ware culture foragers were culturally but not genetically influenced by the Battle Axe culture herders. Am. J. Phys. Anthropol. 172, 638–649 (2020).
Google Scholar
Glob, P. V. Studier over den Jyske Enkeltgravskultur (Gyldendal, 1945).
Müller, J. & Vandkilde, H. in Contrasts of the Nordic Bronze Age. Essays in Honour of Christopher Prescott (eds. Austvoll, K.I., Hem Eriksen, M., Fredriksen, P.D., Melheim, A.L., Prøsch-Danielsen, L., Skogstrand, L.) 29–48 (Brepols, 2020).
Iversen, R. The Transformation of Neolithic Societies. An Eastern Danish Perspective on the 3rd Millennium BC Vol. 88 (Jutland Archaeological Society, 2015).
Damm, C. The Danish Single Grave Culture—ethnic migration or social construction? J. Dan. Archaeol. 10, 199–204 (1991).
Google Scholar
Egfjord, A. F.-H. et al. Genomic steppe ancestry in skeletons from the Neolithic Single Grave Culture in Denmark. PLoS ONE 16, e0244872 (2021).
Google Scholar
Grasgruber, P., Sebera, M., Hrazdíra, E., Cacek, J. & Kalina, T. Major correlates of male height: a study of 105 countries. Econ. Hum. Biol. 21, 172–195 (2016).
Google Scholar
Papac, L. et al. Dynamic changes in genomic and social structures in third millennium bce central Europe. Sci. Adv. 7, eabi6941 (2021).
Google Scholar
Blank, M. Mobility, Subsistence and Mortuary practices. An Interdisciplinary Study of Neolithic and Early Bronze Age Megalithic Populations of Southwestern Sweden. PhD thesis, Univ. of Gothenburg (2021).
Winther Johannsen, J. Late Neolithic expansion. Dan. J. Archaeol. 12, https://doi.org/10.7146/dja.v12i1.132093 (2023).
Pedersen, C. B. et al. The iPSYCH2012 case–cohort sample: new directions for unravelling genetic and environmental architectures of severe mental disorders. Mol. Psychiatry 23, 6–14 (2017).
Google Scholar
Odgaard, B. V. The Holocene vegetation history of northern West Jutland, Denmark. Nord. J. Bot. 14, 546–546 (1994).
Google Scholar
Haak, W. et al. in The Indo-European Puzzle Revisited: Integrating Archaeology, Genetics, and Linguistics (eds Kristiansen, K., Kroonen, G. & Willerslev, E.) 63–80 (2023).
Fischer, A., Gotfredsen, A. B., Meadows, J., Pedersen, L. & Stafford, M. The Rødhals kitchen midden—marine adaptations at the end of the Mesolithic world. J. Archaeol. Sci. 39, 103102 (2021).
Bennike, P. in The Danish Storebælt Since the Ice Age (eds Pedersen, L., Fischer, A. & Aaby, B.) 99–105 (A/S Storebælt Fixed Link, 1997).
Warden, L. et al. Climate induced human demographic and cultural change in northern Europe during the mid-Holocene. Sci Rep. 7, 15251 (2017).
Google Scholar
Krossa, V. R. et al. Regional climate change and the onset of farming in northern Germany and southern Scandinavia. Holocene 27, 1589–1599 (2017).
Google Scholar
Iversen, R. Arrowheads as indicators of interpersonal violence and group identity among the Neolithic Pitted Ware hunters of southwestern Scandinavia. J. Anthropol. Archaeol. 44, 69–86 (2016).
Google Scholar
Lidke, G. Violence in the Single Grave Culture of northern Germany? in Sticks, Stones, and Broken Bones: Neolithic Violence in a European Perspective (eds Schulting, R. J. & Fibiger, L.) 139-150 https://doi.org/10.1093/acprof:osobl/9780199573066.003.0008 (Oxford, 2012).
Schroeder, H. et al. Unraveling ancestry, kinship, and violence in a Late Neolithic mass grave. Proc. Natl Acad. Sci. USA 116, 10705–10710 (2019).
Google Scholar
Rasmussen, S. et al. Early divergent strains of Yersinia pestis in Eurasia 5,000 years ago. Cell 163, 571–582 (2015).
Google Scholar
Rascovan, N. et al. Emergence and spread of basal lineages of Yersinia pestis during the Neolithic decline. Cell 176, 295–305.e10 (2019).
Google Scholar
Hinz, M. et al. in Neolithic Diversities: Perspectives from a Conference in Lund, Sweden 43–51 (Lund Univ., 2015).
Feeser, I., Dörfler, W., Kneisel, J., Hinz, M. & Dreibrodt, S. Human impact and population dynamics in the Neolithic and Bronze Age: multi-proxy evidence from north-western Central Europe. Holocene 29, 1596–1606 (2019).
Google Scholar
Margaryan, A. et al. Population genomics of the Viking world. Nature 585, 390–396 (2020).
Google Scholar
Damgaard, P. B. et al. Improving access to endogenous DNA in ancient bones and teeth. Sci. Rep. 5, 11184 (2015).
Google Scholar
1000 Genomes Project Consortium. A global reference for human genetic variation. Nature 526, 68–74 (2015).
Google Scholar
Maier, R., Flegontov, P., Flegontova, O., Işıldak, U., Changmai, P. & Reich, D. On the limits of fitting complex models of population history to f-statistics. Elife 12, e85492 (2023).
Browning, B. L. & Browning, S. R. Detecting identity by descent and estimating genotype error rates in sequence data. Am. J. Hum. Genet. 93, 840–851 (2013).
Google Scholar
Appadurai, V. et al. Accuracy of haplotype estimation and whole genome imputation affects complex trait analyses in complex biobanks. Commun. Biol. 6, 101 (2023).
Google Scholar
Purcell, S. et al. PLINK: a tool set for whole-genome association and population-based linkage analyses. Am. J. Hum. Genet. 81, 559–575 (2007).
Google Scholar
Walsh, S. et al. The HIrisPlex system for simultaneous prediction of hair and eye colour from DNA. Forensic Sci. Int. Genet. 7, 98–115 (2013).
Google Scholar
Bycroft, C. et al. The UK Biobank resource with deep phenotyping and genomic data. Nature 562, 203–209 (2018).
Google Scholar
Bronk Ramsey, C. Development of the radiocarbon calibration program OxCal. Radiocarbon 43, 355–363 (2001).
Google Scholar
Bronk Ramsey, C. Deposition models for chronological records. Q. Sci. Rev. 27, 42–60 (2008).
Google Scholar
Bronk Ramsey, C. Bayesian analysis of radiocarbon dates. Radiocarbon 51, 337–360 (2009).
Google Scholar
Bronk Ramsey, C. Dealing with outliers and offsets. Radiocarbon 51, 1023–1045 (2009).
Google Scholar
Reimer, P., Austin, W. & Bard, E. The IntCal20 Northern Hemisphere radiocarbon age calibration curve (0–55 cal kbp). Radiocarbon 62, 725–757 (2020).
Google Scholar
Karlsberg, A. J. Flexible Bayesian Methods for Archaeological Dating. PhD thesis, Univ. Sheffield (2006).
Lee, S. & Ramsey, C. Development and application of the trapezoidal model for archaeological chronologies. Radiocarbon 54, 107–122 (2012).
Google Scholar
Meadows, J. et al. Dietary freshwater reservoir effects and the radiocarbon ages of prehistoric human bones from Zvejnieki, Latvia. J. Archaeol. Sci. 6, 678–689 (2016).
Rose, H. A., Meadows, J. & Bjerregaard, M. High-resolution dating of a medieval multiple grave. Radiocarbon 60, 1547–1559 (2018).
Google Scholar
Hedges, R. E. M., Clement, J. G., David, C., Thomas, L. & O’Connell, T. C. Collagen turnover in the adult femoral mid-shaft: modeled from anthropogenic radiocarbon tracer measurements. Am. J. Phys. Anthropol. 133, 808–816 (2007).
Google Scholar
Jørkov, M. L. S., Heinemeier, J. & Lynnerup, N. The petrous bone-a new sampling site for identifying early dietary patterns in stable isotopic studies. Am. J. Phys. Anthropol. 138, 199–209 (2009).
Google Scholar
Schoeninger, M. J. & Moore, K. Bone stable isotope studies in archaeology. J. World Prehist. 6, 247–296 (1992).
Google Scholar
Hedges, R. E. M. & Reynard, L. M. Nitrogen isotopes and the trophic level of humans in archaeology. J. Archaeol. Sci. 34, 1240–1251 (2007).
Google Scholar
Reimer, P. et al. Laboratory protocols used for AMS radiocarbon dating at the 14Chrono Centre. English Heritage Research Report Series 5-2015 https://historicengland.org.uk/research/results/reports/6272/TheQueen%E2%80%99sUniversityBelfast_LaboratoryprotocolsusedforAMSradiocarbondatingatthe14CHRONOCentre (2015).
Longin, R. New method of collagen extraction for radiocarbon dating. Nature 230, 241–242 (1971).
Google Scholar
Ambrose, S. H. & DeNiro, M. J. The isotopic ecology of East African mammals. Oecologia 69, 395–406 (1986).
Google Scholar
van Klinken, G. J. Bone collagen quality indicators for palaeodietary and radiocarbon measurements. J. Archaeol. Sci. 26, 687–695 (1999).
Google Scholar
Alexander Bentley, R. Strontium isotopes from the earth to the archaeological skeleton: A review. J. Archaeol. Method Theory 13, 135–187 (2006).
Google Scholar
Frei, K. M. & Price, T. D. Strontium isotopes and human mobility in prehistoric Denmark. Archaeol. Anthropol. Sci. 4, 103–114 (2012).
Google Scholar
Holt, E., Evans, J. A. & Madgwick, R. Strontium (87Sr/86Sr) mapping: a critical review of methods and approaches. Earth Sci. Rev. 216, 103593 (2021).
Google Scholar
Price, T. D., Burton, J. H. & Bentley, R. A. Characterization of biologically available strontium isotope ratios for the study of prehistoric migration. Archaeometry 44, 117–135 (2002).
Google Scholar
Thomsen, E., Andreasen, R. & Rasmussen, T. L. Homogeneous glacial landscapes can have high local variability of strontium isotope signatures: implications for prehistoric migration studies. Front. Ecol. Evol. 8, 588318 (2021).
Google Scholar
Price, T. D., Klassen, L. & Sjögren, K. G. Pitted ware culture: isotopic evidence for contact between Sweden and Denmark across the Kattegat in the Middle Neolithic, ca. 3000 bc. J. Anthropol. Archaeol. 61, 101254 (2021).
Google Scholar
Hede, M. U. Holocene Climate and Environmental Changes Recorded in High-resolution Lake Sediments from Højby Sø, Denmark. PhD thesis, Univ. Copenhagen (2008).
Sugita, S. Theory of quantitative reconstruction of vegetation I: pollen from large sites REVEALS regional vegetation composition. Holocene 17, 229–241 (2007).
Google Scholar
Sugita, S. Theory of quantitative reconstruction of vegetation II: all you need is LOVE. Holocene 17, 243–257 (2007).
Google Scholar
Nielsen, A. B. et al. Quantitative reconstructions of changes in regional openness in north-central Europe reveal new insights into old questions. Q. Sci. Rev. 47, 131–147 (2012).
Google Scholar
Githumbi, E. et al. Pollen-based maps of past regional vegetation cover in Europe over twelve millennia—evaluation and potential. Front. Ecol. Evol. https://doi.org/10.3389/fevo.2022.795794 (2022).
Nielsen, A. B. & Odgaard, B. V. Quantitative landscape dynamics in Denmark through the last three millennia based on the landscape reconstruction algorithm approach. Veg. Hist. Archaeobot. 19, 375–387 (2010).
Google Scholar
Søe, N. E., Odgaard, B. V., Nielsen, A. B., Olsen, J. & Kristiansen, S. M. Late Holocene landscape development around a Roman Iron Age mass grave, Alken Enge, Denmark. Veg. Hist. Archaeobot. 26, 277–292 (2017).
Google Scholar
Mazier, F. et al. Testing the effect of site selection and parameter setting on REVEALS-model estimates of plant abundance using the Czech Quaternary Palynological Database. Rev. Palaeobot. Palynol. 187, 38–49 (2012).
Google Scholar